This makes me wonder if birds react the same way dogs do in regards to 'pack' mentality...meaning, if the 'lead dog' is not concerned, then the more submissive/anxious dog is less likely to get worked up. Many times, people who have a chronically nervous/scared dog seek out a companion who is more confident to help boost their other dogs courage, so to speak. Also, engaging them in activities such as agility to boost their own confidence in a one-on-one setting.
Is Echo your only bird? ...I'm not saying you should run out and get another, by any means...But in my mind, a bird that instinctually relies on a flock to "lookout" for them, being alone could have an effect on them. It could be why he is more flighty, or scared.
Also, making a big deal out of his fright episodes can sort of reinforce his fear...like rushing to him, coddling, worrying over him, etc. They read your body language so profoundly, that this could worry him further. Also, when it comes to noises like kiss noises and things you *notice* freak him out, try integrating them into benign, good activities...such as making breakfast, building toys, cleaning the cage, etc...
With Doodle, when he takes off in fear of something, I just calmly go pick him up and set him right back where he was prior to his take-off. If he takes off and darts to me for 'comfort' I *still* set him calmly back where he was...and then go get him when he has settled down a bit. I don't know if this makes a difference...lol.